求大侠帮忙解释杨鹏难句中的一个难句

alexzsu

No. 040

But achieving necessary matches in physical properties across interfaces between living and no-living matter requires knowledge of which molecules control the bonding of cells to each other – an area that we have not yet explored thoroughly.

句中的control不应用controling吗？书中的解释of which molecules是用于修饰knowledge，which在此不是定语从句的引导词，那么control不应用controling吗？

buda

还没看到这个句子，但是我怎么觉得 knowledge of which molecules control the bonding of cells to each other 中把of which理解成that, molecules control the bonding of cells to each other 是knowledge 的内容。molecules是复数，自然用control 。不知道是不是这样，