Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
(A) which kept (B) that keeps (C) which has kept (D) that has been keeping (E) having kept