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我觉得你可以这样理解: (Such variations had been demonstrated in nerve cells as in size, shape, chemistry, conduction speed, excitation threshold, and the like) remained negligible in significance for any possible correlation with the manifold dimensions of mental experience.
Variations 是主语 谓语 Remained
原句: (Such variations in size, shape, chemistry, conduction speed, excitation threshold, and the like as had been demonstrated in nerve cells)remained negligible in significance for any possible correlation with the manifold dimensions of mental experience.
中心就是:Variations in xx, xx, xx, and xx such as (those that) had been demonestrated....., remained.............
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