However, this relation is complicated in intertidal dwellers such as commuter diatoms by the fact that these organisms are exposed to the solar-day cycle as well as to the tidal cycle, and sometimes display both solar-day and tidal periods in a single behavior. Commuter diatoms, for example, emerge only during those low tides that occur during the day.
理解的时候可以以这个语序:However, in intertidal dwellers such as commuter diatoms,this relation is complicated by the fact that these organisms (指代commuter diatoms)are exposed to the solar-day cycle as well as to the tidal cycle, and (these organisms) sometimes display both solar-day and tidal periods in a single behavior. For example, commuter diatoms emerge only during those low tides that occur during the day. 尽管如此,在诸如commuter diatoms此类的潮汐住民中,这个关系尤为复杂因为这些生物同时暴露在日光周期和潮汐周期之下,因此这些生物有时会在一个动作表现下同时展现出日光周期和潮汐周期(的特性)。例如,CD只在白天的低潮期才会浮现。