Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
(A) which kept
(B) that keeps
(C) which has kept
(D) that has been keeping
(E) having kept