询问一道语法题

cyx1106

询问一道语法题
Building on civilizations that preceded them in coastal Peru, the Mochica developed their own elaborate society, based on cultivating such crops like corn and beans, the harvesting of fish and shellfish, and exploiting other wild and domestic resources.

A.based on cultivating such crops like corn and beans, the harvesting of fish and shellfish, and exploiting

B.based on the cultivation of such crops as corn and beans, the harvesting of fish and seafood, and the exploitation of  请问正确选项B为什么在BASED 与前面的society加逗号？不是定语从句修饰society吗？
C.and basing it on the cultivation of crops like corn and beans, harvesting fish and seafood, and the exploiting of

D.and they based it on their cultivation of crops such as corn and beans, the harvest of fish and seafood, and exploiting (B)

E.and they based it on their cultivating such crops like corn and beans, their harvest of fish and shellfish, and they exploited