Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
7.Carnivorous mammals can endure what would otherwise be lethal levels of body heat because they have a heat-exchange network which kept the brain from getting too hot.
which kept that keeps
which has kept
that has been keeping having kept
Idiom + Verb form
The two underlined words contain two separate errors. The pronoun which introduces nonrestrictive clauses, which include information relevant but not critical to an understanding of the sentence. The pronoun that introduces a restrictive clause, which is critical to understanding the sentence because the clause defines its antecedent. Here, the clause following which defines heat-exchange network, so that must be used in place of which. The use of the past tense (kept) is incorrect because a current situation is discussed; the present tense (keeps) is consistent with the other verbs in the sentence.
A Which is incorrectly used for a clause that defines; kept is the wrong tense
B Correct. That introduces a restrictive clause; keeps indicates a current situation and is consistent with the other verbs in the sentence.
C Which incorrectly introduces a restrictive clause; has kept is the wrong tense
D The complete sentence is written in the present tense; there is no reason to switch tenses in the restrictive clause
E Having is awkward and imprecise; kept is the incorrect tense