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this should be (e).
the other four choices all misuse "which" (or "each of which", whose modifier properties are the same as those of "which").
in (a), "each of which" doesn't refer to anything at all (it seems to refer to "reserve", but that doesn't make any sense)
in (b), "which" refers to the shark's jaws (this doesn't make sense)
in (c) and (d), "each one of which" refers to the shark's jaws (this doesn't make sense)
also:
(a) and (b) have the wrong meaning: the placement of the "when"/"whenever" clause suggests that the teeth are only there when another tooth is lost or worn down.
(c) has subject-verb disagreement ("each one of which" is singular, but "are" is plural).
(d) is a sentence fragment.
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