逗号加-ing可以做定语吗

mattsun

OG2020 Verbal 分册的题目:

The completion in 1925 of the Holland Tunnel, linking Manhattan with New Jersey's highways, which permitted 2,000 cars to pass through each tube every hour and was hailed as the decade's $48 million engineering masterpiece.
A Tunnel, linking Manhattan with New Jersey's highways, which permitted 2,000 cars to pass through each tube every hour and
B Tunnel, linking Manhattan with New Jersey's highways and permitting 2,000 cars to pass through each tube every hour, it
C Tunnel, linking Manhattan with New Jersey's highways and permitting 2,000 cars to pass through each tube every hour,
D Tunnel linked Manhattan with New Jersey's highways, which permitted 2,000 cars to pass through each tube every hour and
E Tunnel linked Manhattan with New Jersey's highways, permitting 2,000 cars to pass through each tube every hour,


答案选C. 我的问题是关于linking和permitting的语法成分。我记得逗号+ ing只能做状语修饰语，并且修饰的是句子的主语，而不能做定语。但是在这题里面句子主语是completion, 而linking和permitting看上去像是做Tunnel的定语。请问在这里的语法成分是什么?