Both the caribou and the reindeer belong to the species Rangifer tarandus, but after 7,000 years of domestication in Eurasia, reindeer have developed a tendency to circle in tight groups, while caribou tend to spread far and wide
这个选项中的reindeer,谓语动词为什么是have developed,而不是has 呢? 包括后面的caribou tend to,而不是tends to.