35. GWD31-35求助求助求助

liwebber

A consortium of historically Black colleges in the United States capitalizing on such schools traditionally rich relationships with African nations, integrates African concerns into an international business study

A. capitalizing on such school’ traditionally rich relationships with African nations
B. while capitalizing on the traditionally rich relationships of such schools and African nations
C. through capitalizing on such schools’ and African nations’ traditionally rich relationships
D. which capitalize on the traditionally rich relationships of such schools and African nations
E. in capitalizing on such schools’ and African nations’ traditionally rich relationship
这道题的答案是A，我查了以前的求助帖，可是没找到答案，我想问一下，这句话的位于动词是integrates吗？为什么这个谓语动词的前头会有逗号呢？谢谢各位大侠。