求助prep 162

shunwen

Such orb-weaving spiders like the argiope build webs in an essentially wheel-like structure, with an outer rim and a number of spokes emanating from the hub.
A.    Such orb-weaving spiders like the argiope build webs in an essentially wheel-like structure
B.    Such orb-weaving spiders as the argiope builds webs with what is essentially a wheel-like structure
C.    The argiope spider, in the same way as other orb-weaving spiders, build webs in a structure that is essentially wheel-like
D.    D. Orb-weaving spiders such as the argiope build webs that are essentially wheel-like  
E.    Like other orb-weaving spiders do, the argiope builds webs as essentially wheel-like
答案是D，可是我觉得按照D的话，句子就没谓语了。因为那个that are essentially wheel-like 是修饰语，而后面部分又是独立主格。
真是不知道这题怎么回事？
哪位高人可以指点下

hounaiwu

。。。。不是有谓语吗？为什么说没谓语呢？
谓语不是build 吗？？？？

主干 就是  orb-weaving spiders  build webs.