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Mary buys cookies made with the artificial sweetener SugarFree, which tastes as sweet as the corn syrup that her brother loves but which has fewer calories than does an equivalent amount of corn syrup.
前面表红的that 修饰 the corn syrup, 如果后面but 没有which 的话 就可能会被解读成 the corn syrup that her brother loves, but has fewer calories 虽然如果看下去的话这句句子是nonsense 但是加出which 的话就能很明确的表示是在修饰sweetner, 剩下读者的脑细胞了
回到你提出的第二个问题去掉as...as... 这句话就不对了 因为 ,which tastes sweet the corn syrup 句子是不符合语法习惯的 which tastes sweet 才是对的 没有了as...as... the corn syrup 在句子里没有合理的位置
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