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From my opinion, "they" refer to. Images and temples
X was expended on A and on B in which they was enshrined .Obviously, they refer to a and b
From other points, in which could be equal to when
Use when repalce In which ,you could earlier to find out who is the antecedent.
Another opinions from Manhattan Prep: The unclear pronoun in the sentence could be ignored to some degree.
In short, except the disagree of the singular and plural of the antecedent , do not pay much attention on the pronoun in detail.
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