Both the complexity of the phenomenon known as extinction and the vastness of the biosphere, having prompted many scientists to call for a large increase in the number of biologists working both in the field and in laboratories to clarify the relationships among the planet's many endangered life-forms.
Both the complexity of the phenomenon known as extinction and the vastness of the biosphere, having prompted many scientists to call for a large increase in the number of biologists working both in the field and in laboratories to clarify the relationships among the planet's many endangered life-forms.
(A) and the vastness of the biosphere, having prompted many scientists to call for a large increase in the number of biologists working both (B) and the vastness of the biosphere has prompted many scientists calling for a large increase in the number of biologists both working (C) and the vastness of the biosphere have prompted many scientists to call for a large increase in the number of biologists working both (D) as well as the vastness of the biosphere have prompted many scientists to call for a large increase in the number of biologists who both work (E) as well as the vastness of the biosphere has prompted many scientists to call for a large increase in the number of biologists who work both 怎么判断主语是什么呢,是the complexity of the phenomenon known as extinction and the vastness of the biosphere还是the vastness 呢?为什么用的是have而不是has 呢?