[求助]大全496 and平行并列

496.In the first period of religious reformism, Islamic political militancy was manifested in the rise of the Wahhabis in Arabia, the Sanusi in Cyrenaica, the Fulani in Nigeria, and the Mahdi in the Sudan
                        ,  
                    and in  the victory of the Usuli “mujtahids” in Shiite Iran and Iraq.

我知道 the Wahhabis in Arabia, the Sanusi in Cyrenaica, the Fulani in Nigeria, and the Mahdi in the Sudan 是平行并列关系。

was manifested in the rise of  和the victory of 是平行并列关系。

但是为什么会都那个逗号，按道理讲，应该没那个逗号。请NN指点！！