求助gwd-3-3, v+ing 该怎么用？

GWD 3-3

A study by the Ocean Wildlife Campaign urged states to undertake a number of remedies to reverse a decline in the shark population, which includes the establishment of size limits for shark catches, closing state waters for shark fishing during pupping season, and requiring commercial fishers to have federal shark permits.

1. which includes the establishment of size limits for shark catches, closing
2. which includes establishing limits to the size of sharks that can be caught, closing
3. which include the establishment of size limits for shark catches, the closing of
4. including establishing size limits for shark catches, closing
               
5. including the establishing of limits to the size of sharks that are caught, the closing of

including比较特殊, 建议把他就当作介词看

这里是肯定不能用定从的, 因为会有修饰population的歧义

另外B里面的错误还包括把A的shark catches改为sharks that can be caught, 属于错误添加情态动词

B中includes改为include还是修饰population啊？我也是对分词的修饰大感困惑！