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I've expanded the rule a bit:
When an introductory modifier includes a pronoun without an antecedent, the referent should be the FIRST WORD OF THE FOLLOWING CLAUSE.
In 99% of cases, the first word of the following clause will be the SUBJECT.
The SC above presents an exception.
In the OA here (D), the referent for her (possessive pronoun) is another possessive construction (Mary Leakey's).
It is perfectly fine for a possessive pronoun to serve to refer to another possessive construction.
Since each of these constructions serves as an ADJECTIVE, the first word of the following clause is not the subject but an adjective.
The introductory modifier here serves as an ADVERB describing how Mary Leakey's contributions INCLUDE.
Mary Leakey's contributions INCLUDE -- in addition to her work on the Miocene hominid fossil record -- her discovery of the earliest direct evidence of hominid activity and her painstaking documentation of East African cave paintings.
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