Prep_SC1_052迷惑，求助，请指点……

In addition to her work on the Miocene hominid
fossil record, Mary Leakey contributed to archaeology with her discovery of
the earliest direct evidence of hominid activity and painstakingly documenting
East African cave paintings.

(A) Leakey contributed to archaeology with her
discovery of the earliest direct evidence of hominid activity and painstakingly
documenting

(B) Leakey contributed to archaeology by her
discovery of the earliest direct evidence of hominid activity and by
painstakingly documenting

(C) Leakey was a contributor to archaeology with
her discovery of the earliest direct evidence of hominid activity and with her
painstaking documentation of

(D) Leakey's contributions to archaeology include
her discovery of the earliest direct evidence of hominid activity and her
painstaking documentation of

(E) Leakey's contributions to archaeology include
discovering the earliest direct evidence of hominid activity and painstaking
documentation of

正确答案是：D，请问如果正确答案是D，那么这句话的谓语是什么？