prep 123 请教

123. As a result of a supernova explosion, every human being on Earth was bombarded on February 23, 1987, by about 100 billion neutrinos; fortunately, neutrinos are harmless elementary particles that are produced in nuclear reactions and that interact very weakly with matter.

A. neutrinos are harmless elementary particles that are produced in nuclear reactions and that

B. neutrinos, which are harmless, are elementary particles produced in nuclear reactions and which

C. neutrinos are harmless elementary particles produced in nuclear reactions and which

D. these harmless elementary particles are produced in nuclear reactions, and neutrinos

E. these elementary particles, harmless products of nuclear reactions, are neutrinos that

B: and which 没有指代对象

C: 同上

D: these无指代

E: 同上

注意：

1、that … and that … 做限定性定语从句的并列,   , which and which 比较少用
                            

为什么E不对呢，these无指代，前文不是有一个 neutrinos么？？？