主谓搭配问题！！！！！

When an active tooth in the shark’s jaws is lost or worn down, many spare teeth lie in seemingly limitless reserve, each of which are ready to slide into the appropriate position.

1. When an active tooth in the shark’s jaws is lost or worn down, many spare teeth lie in seemingly limitless reserve, each of which are ready to slide into the appropriate position.
2. Whenever an active tooth is lost or worn down, many spare teeth lie in seemingly limitless reserve in the shark’s jaws, which are each ready to slide into the appropriate position.
3. Many spare teeth lie in seemingly limitless reserve in the shark’s jaws, each one of which are ready to slide into the appropriate position when an active tooth is lost or worn down.
4. The many spare teeth lying in seemingly limitless reserve in the shark’s jaws, each one of which is ready to slide into the appropriate position whenever an active tooth is lost or worn down.
5. In the shark’s jaws, many spare teeth lie in seemingly limitless reserve, each one ready to slide into the appropriate position whenever an active tooth is lost or worn down.
                       

GWD 给的答案是A，但是这里主谓搭配有问题：each后面怎么是are而不是is??? 

我选的答案是：D，但是好像each one 是语义重复

请求帮助

没人帮忙，自己顶一下