OG120 的小疑问

OG120.

The Parthenon was a church from 1204 until 1456, when Athens was taken by General Mohammed the Conqueror, the Turkish sultan, who established a mosque in the building and used the Acropolis as a fortress.
(A) who established a mosque in the building and used the Acropolis as
(B) who, establishing a mosque in the building, used the Acropolis like
(C) who, when he had established a mosque in the building, used the Acropolis like
(D) who had established a mosque in the building, using the Acropolis to be
(E) establishing a mosque in the building and using the Acropolis as

对这题的答案我没有意见，但E选项的解释使我产生了一个疑问，那就是：“－ING”形式作修饰语时到底是就近修饰还是修饰主句主语，事实上新东方老师是说就近修饰，但E的解释无疑是在说修饰主语，而我两者的情况都见过，都被搞晕了>.<