GWD-24-34

A.    Studying the fruit fly, a household nuisance but a time-honoredexperimental subject, has enabled the secrets of how embryos develop to beginto be unraveled by scientists.

B.    By the study of the fruit fly, a household nuisance and also atime-honored experimental subject, it was possible for the secrets of howembryos develop to begin to be unraveled by scientists.

C.    By studying a household nuisance but a time-honored experimentalsubject, the fruit fly enabled scientists to begin to unravel the secrets ofhow embryos develop.

D.     By studying the fruitfly, a household nuisance and also a time-honored experimental subject, thesecrets of how embryos develop are beginning to be unraveled byscientists.     (E)

E.    The study of the fruit fly, a household nuisance but a time-honoredexperimental subject, has enabled scientists to begin to unravel the secrets ofhow embryos develop.

这道题的正确答案是E，但我有点不明白The study of the fruit fly, a household nuisance but a time-honored experimental subject      a household nuisance..虽然语义上没有问题，但在语法上.作为同位语难道没有修饰the study的歧义吗，A OF B结构同位语可以不管A只修饰B吗

另外，我觉得D特别好，看了语法笔记的答案还是无法说服我自己不选D，求NN指教，跪求！！！！！！！！