Around 1900, fishermen inthe Chesapeake Bay area landed more than seventeen million pounds of shad in asingle year, but by 1920, overfishing and the proliferation of milldams andculverts that have blocked shad migrations up their spawning streams hadreduced landings to less than four million pounds.
A. that have blocked shadmigrations up their spawning streams had reduced landings to less
B. that blocked shad frommigrating up their spawning streams had reduced landings to less
C. that blocked shad frommigrating up their spawning streams reduced landings to a lower amount
D. having blocked shadfrom migrating up their spawning streams reduced landings to less
E. having blocked shadmigrations up their spawning streams had reduced landings to an amount lower
请教NN们 that blocked shad from migrating up their spawning streams 与having blocked shad from migrating up their spawning streams 的区别,主要是这里为什么一定要用that从句修饰,而不能用V-ing修饰milldams and culverts 呢?是因为时间点不同吗?谢谢
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