49. As an actress and, more importantly, as a teacher of acting, Stella Adler was one of the most influential artists in the American theater, who rained several generations of actors including Marlon Brando and Robert De Niro.
(A) Stella Adler was one of the most influential artists in the American theater, who trained several generations of actors including
(B) Stella Adler, one of the most influential artists in the American theater, trained several generations of actors who include
(C) Stella Adler was one of the most influential artists in the American theater, training several generations of actors whose ranks included
(D) one of the most influential artists in the American theater was Stella Adler, who trained several generations of actors including （C）
(E) one of the most influential artists in the American theater, Stella Adler, trained several generations of actors whose ranks included
这道题B为什么不对？答案是C，但C里面我加亮部分是什么意思？
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