The work of mathematician Roger Penrose in the early 1970s, on the geometry of what are called aperiodic tiles, turned out to describe the architecture of a previously unknown class of crystals.
(A) what are called aperiodic tiles, turned out to describe
(B) what is called aperiodic tiles, describes
(C) aperiodic tiles, describing
(D) so-called aperiodic tiles, describe（A）
(E) aperiodic tiles, it turned out to describe
on the geometry of what are called aperiodic tiles在句子中做什么成分？是定语从句修饰work? 但是如果是定语从句先行词单数work，从句谓语动词怎么又是were?
晕ing!!!!!!!!!!!
help!
作插入语
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