请教885-9-27

27. The Parthenon was a church from 1204 until 1456, when Athens was taken by General Mohammed the Conqueror, the Turkish sultan, who established a mosque in the building and used the Acropolis as a fortress.

(A) who established a mosque in the building and used the Acropolis as

(B) who, establishing a mosque in the building, used the Acropolis like

(C) who, when he had established a mosque in the building, used the Acropolis like

(D) who had established a mosque in the building, using the Acropolis to be

(E) establishing a mosque in the building and using the Acropolis as

答案是A, 为什么E不可以，分词做定语修饰不是比从句更好吗？而且此处并不存在一次和反复的问题啊

分词做定语是好，但不是最好，它与定语从句相比，我认为在这里是都可用的，

但E中分词前是逗号啊，是否作成状语形式了，去逗号，对