Fossils of a whale that beached on an African shore more than a million years ago and was subsequently butchered by hominids have been recovered by paleontologists.
A. that beached on an African shore more than a million years ago and was subsequently butchered by hominids have
B. that beached on an African shore more than a million years ago and then was subsequently butchered by hominids has
C. that beached on an African shore more than a million years ago, which was subsequently butchered by hominids, has
D. having been beached on an African shore more than a million years ago and subsequently butchered by hominids, have
E. having beached on an African shore more than a million years ago and then subsequently were butchered by hominids have