大家好，
可否指点下题。此题答案A, 但是本人感到全句根本就没有谓语，不完整，如何为句子。征求各位高见？谢谢。
Salt deposits and moisture threaten to destroy the Mohenjo-Daro excavation in Pakistan, the site of an ancient civilization that flourished at the same time as the civilizations in the Nile delta and the river valleys of the Tigris and Euphrates.
(A) that flourished at the same time as the civilizations
(B) that had flourished at the same time as had the civilizations
(C) that flourished at the same time those had
(D) flourishing at the same time as those did
(E) flourishing at the same time as those were
threaten
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